Bussing, W. A., and M. I. López. “Fishes Collected During the Victor Hensen Costa Rica Expedition (1993/1994)”. Revista De Biología Tropical, vol. 44, no. S3, Dec. 1996, pp. 183–186, https://www.revistas.ucr.ac.cr/index.php/rbt/article/view/29564.